In this section, you will explore the following questions:

  • What are the two major abiotic factors that determine terrestrial biomes?
  • What are distinguishing characteristics of each of the major terrestrial biomes?

Connection for AP® Courses

Much of the information in this section is outside the scope of AP®. You do not need to memorize a list of Earth’s major terrestrial biomes and their descriptive features. However, we learned previously that organisms interact with each other and with their environments to move matter and energy. Biomes are ripe with examples of these interactions. A biome refers to a major type of terrestrial (or aquatic) community distributed according to climate, which determines the predominant vegetation. In turn, the vegetation influences what types of animals can inhabit the area. Comparing the annual totals of precipitation and fluctuations in precipitation from one biome to another provides clues as to the importance of abiotic factors in the distribution of biomes. The same type of biome can occur in different areas of the world (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12).

Information presented and the examples highlighted in the section support concepts outlined in Big Idea 2 of the AP® Biology Curriculum Framework. The AP® Learning Objectives listed in the Curriculum Framework provide a transparent foundation for the AP®Biology course, an inquiry-based laboratory experience, instructional activities, and AP® exam questions. A learning objective merges required content with one or more of the seven science practices.

Big Idea 2 Biological systems utilize free energy and molecular building blocks to grow, to reproduce, and to maintain dynamic homeostasis.
Enduring Understanding 2.D Growth and dynamic homeostasis of a biological system are influenced by changes in the system’s environment.
Essential Knowledge 2.D.1 Abiotic factors in an environment determine the characteristics of the environment (biome).
Science Practice 1.3 The student can refine representations and models of natural or man-made phenomena and systems in the domain.
Science Practice 3.2 The student can refine scientific questions.
Learning Objective 2.22 The student is able to refine scientific models and questions about the effect of complex biotic and abiotic interactions on biological systems, from cells and organisms to populations, communities, and ecosystems.
Essential Knowledge 2.D.1 Abiotic factors in an environment determine the characteristics of the environment (biome).
Science Practice 5.1 The student can analyze data to identify patterns or relationships.
Learning Objective 2.24 The student is able to analyze data to identify possible patterns and relationships between a biotic or abiotic factor and a biological system (cells, organisms, populations, communities, or ecosystems).

VISUAL CONNECTION

 This world map shows the eight major biomes, polar ice, and mountains. Tropical forests, deserts and savannas are found primarily in South America, Africa, and Australia. Tropical forests also dominate Southeast Asia. Deserts dominate the Middle East and are found in the southwestern United States. Temperate forests dominate the eastern United States, Europe, and Eastern Asia. Temperate grasslands dominate the midwestern United States and parts of Asia, and are also found in South America. The boreal forest is found in northern Canada, Europe, and Asia, and tundra exists to the north of the boreal forest. Mountainous regions run the length of North and South America, and are found in northern India, Africa, and parts of Europe. Polar ice covers Greenland and Antarctica, which the latter is not shown on the map.
Figure 35.12Each of the world’s major biomes is distinguished by characteristic temperatures and amounts of precipitation. Polar ice and mountains are also shown.
Which of the following statements about biomes is false?
  1. Chapparal is dominated by shrubs.
  2. Grasses dominate savannas and temperate grasslands.
  3. Boreal forests are dominated by deciduous trees.
  4. Lichens are common in the arctic tundra.

Tropical Wet Forest

Tropical wet forests are also referred to as tropical rainforests. This biome is found in equatorial regions (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). The vegetation is characterized by plants with broad leaves that fall off throughout the year. Unlike the trees of deciduous forests, the trees in this biome do not have a seasonal loss of leaves associated with variations in temperature and sunlight; these forests are “evergreen” year-round.

The temperature and sunlight profiles of tropical wet forests are very stable in comparison to that of other terrestrial biomes, with the temperatures ranging from 20 °C to 34 °C (68 °F to 93 °F). When one compares the annual temperature variation of tropical wet forests with that of other forest biomes, the lack of seasonal temperature variation in the tropical wet forest becomes apparent. This lack of seasonality leads to year-round plant growth, rather than the seasonal (spring, summer, and fall) growth seen in other biomes. In contrast to other ecosystems, tropical ecosystems do not have long days and short days during the yearly cycle. Instead, a constant daily amount of sunlight (11–12 hrs per day) provides more solar radiation, thereby, a longer period of time for plant growth.

The annual rainfall in tropical wet forests ranges from 125 to 660 cm (50–200 in) with some monthly variation. While sunlight and temperature remain fairly consistent, annual rainfall is highly variable. Tropical wet forests have wet months in which there can be more than 30 cm (11–12 in) of precipitation, as well as dry months in which there are fewer than 10 cm (3.5 in) of rainfall. However, the driest month of a tropical wet forest still exceeds the annual rainfall of some other biomes, such as deserts.

Tropical wet forests have high net primary productivity because the annual temperatures and precipitation values in these areas are ideal for plant growth. Therefore, the extensive biomass present in the tropical wet forest leads to plant communities with very high species diversities (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_02">Figure 35.13). Tropical wet forests have more species of trees than any other biome; on average between 100 and 300 species of trees are present in a single hectare (2.5 acres) of South America. One way to visualize this is to compare the distinctive horizontal layers within the tropical wet forest biome. On the forest floor is a sparse layer of plants and decaying plant matter. Above that is an understory of short shrubby foliage. A layer of trees rises above this understory and is topped by a closed upper canopy—the uppermost overhead layer of branches and leaves. Some additional trees emerge through this closed upper canopy. These layers provide diverse and complex habitats for the variety of plants, fungi, animals, and other organisms within the tropical wet forests. For instance, epiphytes are plants that grow on other plants, which typically are not harmed. Epiphytes are found throughout tropical wet forest biomes. Many species of animals use the variety of plants and the complex structure of the tropical wet forests for food and shelter. Some organisms live several meters above ground and have adapted to this arboreal lifestyle.

 This photo depicts a section of the Amazon River, which is brown with mud. Trees line the edge of the river.
Figure 35.13Tropical wet forests, such as these forests of Madre de Dios, Peru, near the Amazon River, have high species diversity. (credit: Roosevelt Garcia)

Savannas

Savannas are grasslands with scattered trees, and they are located in Africa, South America, and northern Australia (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). Savannas are hot, tropical areas with temperatures averaging from 24 °C to 29 °C (75 °F to 84 °F) and an annual rainfall of 10–40 cm (3.9–15.7 in). Savannas have an extensive dry season; for this reason, forest trees do not grow as well as they do in the tropical wet forest (or other forest biomes). As a result, within the grasses and forbs (herbaceous flowering plants) that dominate the savanna, there are relatively few trees (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_03">Figure 35.14). Since fire is an important source of disturbance in this biome, plants have evolved well-developed root systems that allow them to quickly re-sprout after a fire.

 A grassy slope plain is dotted with small trees.
Figure 35.14Savannas, like this one in Taita Hills Wildlife Sanctuary in Kenya, are dominated by grasses. (credit: Christopher T. Cooper)

Subtropical Deserts

Subtropical deserts exist between 15 ° and 30 ° north and south latitude and are centered on the Tropics of Cancer and Capricorn (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). This biome is very dry; in some years, evaporation exceeds precipitation. Subtropical hot deserts can have daytime soil surface temperatures above 60 °C (140 °F) and nighttime temperatures approaching 0 °C (32 °F). In cold deserts, temperatures can be as high as 25 °C and can drop below -30 °C (-22 °F). Subtropical deserts are characterized by low annual precipitation of fewer than 30 cm (12 in) with little monthly variation and lack of predictability in rainfall. In some cases, the annual rainfall can be as low as 2 cm (0.8 in) in subtropical deserts located in central Australia (“the Outback”) and northern Africa.

The vegetation and low animal diversity of this biome is closely related to this low and unpredictable precipitation. Very dry deserts lack perennial vegetation that lives from one year to the next; instead, many plants are annuals that grow quickly and reproduce when rainfall does occur, then they die. Many other plants in these areas are characterized by having a number of adaptations that conserve water, such as deep roots, reduced foliage, and water-storing stems (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_04">Figure 35.15). Seed plants in the desert produce seeds that can be in dormancy for extended periods between rains. Adaptations in desert animals include nocturnal behavior and burrowing.

 This photo shows a sandy desert dotted with scrubby bushes. An ocotillo plant dominates the picture. It has long, thin unbranched stems that grow straight up from the base of the plant and radiate out slightly. The plant has no leaves.
Figure 35.15To reduce water loss, many desert plants have tiny leaves or no leaves at all. The leaves of ocotillo (Fouquieria splendens), shown here in the Sonora Desert near Gila Bend, Arizona, appear only after rainfall, and then are shed.

Chaparral

The chaparral is also called the scrub forest and is found in California, along the Mediterranean Sea, and along the southern coast of Australia (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). The annual rainfall in this biome ranges from 65 cm to 75 cm (25.6–29.5 in), and the majority of the rain falls in the winter. Summers are very dry and many chaparral plants are dormant during the summertime. The chaparral vegetation, shown in //cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_05">Figure 35.16, is dominated by shrubs and is adapted to periodic fires, with some plants producing seeds that only germinate after a hot fire. The ashes left behind after a fire are rich in nutrients like nitrogen that fertilize the soil and promote plant regrowth.

 Photo depicts a landscape with many shrubs, dormant grass, a few trees, and mountains in the background.
Figure 35.16The chaparral is dominated by shrubs. (credit: Miguel Vieira)

Temperate Grasslands

Temperate grasslands are found throughout central North America, where they are also known as prairies; they are also in Eurasia, where they are known as steppes (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). Temperate grasslands have pronounced annual fluctuations in temperature with hot summers and cold winters. The annual temperature variation produces specific growing seasons for plants. Plant growth is possible when temperatures are warm enough to sustain plant growth and when ample water is available, which occurs in the spring, summer, and fall. During much of the winter, temperatures are low, and water, which is stored in the form of ice, is not available for plant growth.

Annual precipitation ranges from 25 cm to 75 cm (9.8–29.5 in). Because of relatively lower annual precipitation in temperate grasslands, there are few trees except for those found growing along rivers or streams. The dominant vegetation tends to consist of grasses and some prairies sustain populations of grazing animals //cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_06">Figure 35.17. The vegetation is very dense and the soils are fertile because the subsurface of the soil is packed with the roots and rhizomes (underground stems) of these grasses. The roots and rhizomes act to anchor plants into the ground and replenish the organic material (humus) in the soil when they die and decay.

 This photo shows a bison, which is dark brown in color with an even darker head. The hind part of the animal has short fur, and the front of the animal has longer, curly fur.
Figure 35.17The American bison (Bison bison), more commonly called the buffalo, is a grazing mammal that once populated American prairies in huge numbers. (credit: Jack Dykinga, USDA Agricultural Research Service)

Fires, mainly caused by lightning, are a natural disturbance in temperate grasslands. When fire is suppressed in temperate grasslands, the vegetation eventually converts to scrub and dense forests. Often, the restoration or management of temperate grasslands requires the use of controlled burns to suppress the growth of trees and maintain the grasses.

Temperate Forests

Temperate forests are the most common biome in eastern North America, Western Europe, Eastern Asia, Chile, and New Zealand (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). This biome is found throughout mid-latitude regions. Temperatures range between -30 °C and 30 °C (-22 °F to 86 °F) and drop to below freezing on an annual basis. These temperatures mean that temperate forests have defined growing seasons during the spring, summer, and early fall. Precipitation is relatively constant throughout the year and ranges between 75 cm and 150 cm (29.5–59 in).

Because of the moderate annual rainfall and temperatures, deciduous trees are the dominant plant in this biome (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_07">Figure 35.18). Deciduous trees lose their leaves each fall and remain leafless in the winter. Thus, no photosynthesis occurs in the deciduous trees during the dormant winter period. Each spring, new leaves appear as the temperature increases. Because of the dormant period, the net primary productivity of temperate forests is less than that of tropical wet forests. In addition, temperate forests show less diversity of tree species than tropical wet forest biomes.

 Photo shows a deciduous forest with many tall trees, some smaller trees and grass, and lots of dead leaves on the forest floor. Sunlight filters down to the forest floor.
Figure 35.18Deciduous trees are the dominant plant in the temperate forest. (credit: Oliver Herold)

The trees of the temperate forests leaf out and shade much of the ground; however, this biome is more open than tropical wet forests because trees in the temperate forests do not grow as tall as the trees in tropical wet forests. The soils of the temperate forests are rich in inorganic and organic nutrients. This is due to the thick layer of leaf litter on forest floors. As this leaf litter decays, nutrients are returned to the soil. The leaf litter also protects soil from erosion, insulates the ground, and provides habitats for invertebrates (such as the pill bug or roly-poly, Armadillidium vulgare) and their predators, such as the red-backed salamander (Plethodon cinereus).

Boreal Forests

The boreal forest, also known as taiga or coniferous forest, is found south of the Arctic Circle and across most of Canada, Alaska, Russia, and northern Europe (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). This biome has cold, dry winters and short, cool, wet summers. The annual precipitation is from 40 cm to 100 cm (15.7–39 in) and usually takes the form of snow. Little evaporation occurs because of the cold temperatures.

The long and cold winters in the boreal forest have led to the predominance of cold-tolerant cone-bearing plants. These are evergreen coniferous trees like pines, spruce, and fir, which retain their needle-shaped leaves year-round. Evergreen trees can photosynthesize earlier in the spring than deciduous trees because less energy from the sun is required to warm a needle-like leaf than a broad leaf. This benefits evergreen trees, which grow faster than deciduous trees in the boreal forest. In addition, soils in boreal forest regions tend to be acidic with little available nitrogen. Leaves are a nitrogen-rich structure and deciduous trees must produce a new set of these nitrogen-rich structures each year. Therefore, coniferous trees that retain nitrogen-rich needles may have a competitive advantage over the broad-leafed deciduous trees.

The net primary productivity of boreal forests is lower than that of temperate forests and tropical wet forests. The aboveground biomass of boreal forests is high because these slow-growing tree species are long lived and accumulate standing biomass over time. Plant species diversity is less than that seen in temperate forests and tropical wet forests. Boreal forests lack the pronounced elements of the layered forest structure seen in tropical wet forests. The structure of a boreal forest is often only a tree layer and a ground layer (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_08">Figure 35.19). When conifer needles are dropped, they decompose more slowly than broad leaves; therefore, fewer nutrients are returned to the soil to fuel plant growth.

 The photo shows a boreal forest with a uniform low layer of plants and tall conifers scattered throughout the landscape. The snowcapped mountains of the Alaska Range are in the background.
Figure 35.19The boreal forest (taiga) has low lying plants and conifer trees. (credit: L.B. Brubaker)

Arctic Tundra

The Arctic tundra lies north of the subarctic boreal forest and is located throughout the Arctic regions of the northern hemisphere (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_01">Figure 35.12). The average winter temperature is -34 °C (-29.2 °F) and the average summer temperature is from 3 °C to 12 °C (37 °F–52 °F). Plants in the arctic tundra have a very short growing season of approximately 10–12 weeks. However, during this time, there are almost 24 hours of daylight and plant growth is rapid. The annual precipitation of the Arctic tundra is very low with little annual variation in precipitation. And, as in the boreal forests, there is little evaporation due to the cold temperatures.

Plants in the Arctic tundra are generally low to the ground (//cnx.org/contents/This email address is being protected from spambots. You need JavaScript enabled to view it.:gvsrptMX@11/35-3-Terrestrial-Biomes#fig-ch44_03_09">Figure 35.20). There is little species diversity, low net primary productivity, and low aboveground biomass. The soils of the Arctic tundra may remain in a perennially frozen state referred to as permafrost. The permafrost makes it impossible for roots to penetrate deep into the soil and slows the decay of organic matter, which inhibits the release of nutrients from organic matter. During the growing season, the ground of the Arctic tundra can be completely covered with plants or lichens.

 This photo shows a flat plain covered with shrub. Many of the shrubs are covered in pink flowers.
Figure 35.20Low-growing plants such as shrub willow dominate the tundra landscape, shown here in the Arctic National Wildlife Refuge. (credit: USFWS Arctic National Wildlife Refuge)

LINK TO LEARNING

Watch this Assignment Discovery: Biomes video for an overview of biomes. To explore further, select one of the biomes on the extended playlist: desert, savanna, temperate forest, temperate grassland, tropic, tundra.

Which of the following biomes is characterized by low precipitation, cactus, hardy shrubs, and high temperatures?
  1. tundra
  2. chapparal
  3. desert
  4. boreal forest

SCIENCE PRACTICE CONNECTION FOR AP® COURSES

ACTIVITY

For five different terrestrial biomes, create a visual representation to describe each biome and factors that affect its climate. What unique adaptations help plants and animals survive in each biome?

THINK ABOUT IT

In what ways are the subtropical desert and the Arctic tundra similar?